Surah An Nahl Tafseer

وَلِلّهِ يَسْجُدُ مَا فِي السَّمَاوَاتِ وَمَا فِي الَارْضِ مِن دَابَّةٍ

And to Allah prostrate all that are in the heavens and all that are in the earth, of the moving creatures,

As Allah says:

وَللَّهِ يَسْجُدُ مَن فِى السَّمَـوَتِ وَالاٌّرْضِ طَوْعًا وَكَرْهًا وَظِلَـلُهُم بِالْغُدُوِّ وَالاٌّصَالِ

And to Allah (alone) all who are in the heavens and the earth fall in prostration, willingly or unwillingly, and so do their shadows in the mornings and in the afternoons. (13:15)

وَالْمَليِكَةُ وَهُمْ لَا يَسْتَكْبِرُونَ

and the angels, and they are not proud.

means, they prostrate to Allah and are not too proud to worship Him